GSD Species Synonymy
Current Name: Arthonia atra (Pers.) A. Schneid., Guide Study Lich.: 131 (1898) Synonymy: Opegrapha atra Pers., Ann. Bot. (Usteri) 7: 30 (1794) Lichen denigratus Ach., Lich. suec. prodr. (Linköping): 24 (1799) Opegrapha denigrata var. atra (Pers.) Ach., Lich. Univ.: 260 (1810) Graphis atra (Pers.) Spreng., Syst. veg., Edn 16 4(1): 251 (1827) Graphis varia var. atra (Pers.) Branth & Rostr., Bot. Tidsskr. 3: 243 (1869) Graphis scripta var. atra (Pers.) Galløe, Bot. Tidsskr. 28: 330, tab. X, fig. 50 (1908)
Opegrapha denigrata (Ach.) Ach., Methodus, Sectio prior (Stockholmiæ): 27 (1803) Opegrapha atra f. denigrata (Ach.) Moug., Bull. Soc. bot. Fr. 4(4): 259 (1857)
Opegrapha stenocarpa var. denigrata (Ach.) Ach., Syn. meth. lich. (Lund): 75 (1814) Hysterina denigrata (Ach.) Gray, Nat. Arr. Brit. Pl. (London) 1: 507 (1821) Opegrapha atra var. denigrata (Ach.) Schaer., Lich. helv. spicil. 1: 48 (1823) Opegrapha atra * denigrata (Ach.) Trevis., Lichenoth. Veneta: no. 199 (1869)
Opegrapha rimosa DC., in Lamarck & de Candolle, Fl. franç., Edn 3 (Paris) 2: 312 (1805) Opegrapha subpulveracea var. rimosa (DC.) L. Marchand, in Hall, Vrolik & Mulder, Bijdr. Natuurk. Wetensch. 4: 279 (1829) Opegrapha atra var. rimosa (DC.) Zahlbr., Cat. Lich. Univers. 2: 174 (1923) Sphinctrina meridionalis var. rimosa (DC.) Zahlbr., Cat. Lich. Univers. 2: 174 (1923) Opegrapha atra f. rimosa (DC.) J. Nowak, Flora Polska, Porosty (Lichenes) (Warszawa-Kraków) 1(1): 126 (1983)
Opegrapha stenocarpa Ach., K. Vetensk-Acad. Nya Handl. 30(2): 100 (1809) Opegrapha atra var. stenocarpa (Ach.) Dufour, J. Phys. Chim. Hist. nat. Arts 87: 221 (1818) Graphis stenocarpa (Ach.) Spreng., Syst. veg., Edn 16 4(1): 250 (1827) Graphis scripta var. stenocarpa (Ach.) Fürnr., Naturhist. Topogr. Regensburg (Regensburg) 2: 241 (1839) Opegrapha vulgata var. stenocarpa (Ach.) Leight., Ann. Mag. nat. Hist., Ser. 2 13(no. 76): 209 (1854) Opegrapha vulgata f. stenocarpa (Ach.) Leight., Ann. Mag. nat. Hist., Ser. 2 13(no. 76): 209 (1854) Opegrapha atra B stenocarpa (Ach.) Trevis., Lichenoth. Veneta 7-8: no. 200 (1869) Opegrapha atra f. stenocarpa (Ach.) J. Nowak, Flora Polska, Porosty (Lichenes) (Warszawa-Kraków) 1(1): 127 (1983)
Opegrapha stenocarpa var. hapalea Ach., Lich. Univ.: 257 (1810) Opegrapha hapalea (Ach.) Ach., Syn. meth. lich. (Lund): 79 (1814) Opegrapha atra var. hapalea (Ach.) Nyl., Mém. Soc. Imp. Sci. Nat. Cherbourg 3: 188 (1855) Opegrapha atra f. hapalea (Ach.) Nyl., Florule Finistère (Paris): 88 (1867)
Opegrapha nimbosa Sm., in Smith & Sowerby, Engl. Bot. 33: [2346] (1812) Hysterina nimbosa Gray, Nat. Arr. Brit. Pl. (London) 1: 504 (1821)
Opegrapha cerasi Chevall., J. Phys. Chim. Hist. nat. Arts 94: 38 (1822) Opegrapha atra f. cerasi Arnold, Flora, Regensburg 67(34): 662 (1884)
Graphis atra var. gyrograpta Wallr., Fl. crypt. Germ. (Norimbergae) 1: 326 (1831)
Opegrapha atra var. parallela Leight. ex Mudd, Man. Brit. Lich.: 232 (1854) Opegrapha atra b parallela (Leight. ex Mudd) Mudd, Man. Brit. Lich.: 232 (1861) Opegrapha atra f. parallela (Leight. ex Mudd) Arnold, Flora, Regensburg 44(42): 660 (1861)
Opegrapha atra var. arthonioidea Leight. ex Mudd, Man. Brit. Lich.: 232 (1861) Opegrapha atra f. arthonioidea (Leight. ex Mudd) Arnold, Flora, Regensburg 44(42): 660 (1861)
Opegrapha atra f. nigrita Leight., in Mudd, Man. Brit. Lich.: 232 (1861) Opegrapha atra f. nigrita (Leight.) Arnold, Flora, Regensburg 44(42): 660 (1861)
Opegrapha atra var. nigrita Leight. ex Mudd, Man. Brit. Lich.: 232 (1861)
Opegrapha atra f. tenuior Nyl. ex Stizenb., Flora, Regensburg 48(5): 74 (1865) Opegrapha calcarea f. tenuior (Nyl. ex Stizenb.) Zahlbr., Cat. Lich. Univers. 2: 181 (1923)
Position in classification: Arthoniaceae, Arthoniales, Arthoniomycetidae, Arthoniomycetes, Ascomycota, Fungi Synonymy Contributor(s): Kew Mycology (2021)
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